Keratins are a family of fibrous structural proteins; tough and insoluble, they form the hard but mineralized structures found in reptiles, birds, amphibians, and mammals. They are rivaled as biological materials in toughness only by chitin.
Keratins are intermediate filaments. Like all intermediate filaments, keratin proteins form filamentous polymers in a series of assembly steps beginning with dimerization; dimers assemble into tetramers and octamers and eventually, the current hypothesis holds, into unit-length-filaments (ULF) capable of annealing end-to-end into long filaments.
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